Classification and Systematics of the Ichneumonidae (Hymenoptera) David Wahl American Entomological Institute

OPHIONINAE

• Ophionidae Shuckard, 1840. Type-genus: Ophion Fabricius.
• Enicospilini Townes, 1971. Type-genus: Enicospilus Stephens.

1. Enicospilus genus-group
   1. Orientospilus subgroup
      1. Orientospilus Morley, 1912; Ethiopian, Oriental
      2. Prethophion Townes, 1971; Neotropical
      3. Simophion Cushman, 1947; Holarctic
   2. Ophiogastrella subgroup
      1. Ophiogastrella Brues, 1912; Neotropical

         Brachyscenia Enderlein, 1921

   3. Stauropoctonus subgroup
      1. Lepiscelus Townes, 1971; Ethiopian
      2. Stauropoctonus Brauns, 1889; worldwide except Nearctic

         Stauropodoctonus Morley, 1913

         Nipponophion Uchida, 1928

         Aulophion Cushman, 1947 (Gauld, 1985)

   4. Leptophion subgroup
      1. Laticoleus Townes, 1973; Ethiopian
      2. Leptophion Cameron, 1901; Australian, Oriental

         Spilophion Cameron, 1905

         Coiloneura Szépligeti, 1905

      3. Pamophion Gauld, 1977a; Australian
   5. Enicospilus subgroup
      1. Abanchogastra Perkins, 1902; Hawaii (Gauld, 1985)
      2. Banchogastra Ashmead, 1900; Hawaii
      3. Dicamptus Szépligeti, 1905; Australian, Ethiopian, Oriental, Palearctic
      4. Enicospilus Stephens, 1835; worldwide

         Henicospilus Agasssiz, 1846

         Allocamptus Förster, 1869

         Dispilus Kriechbaumer, 1894

         Pleuroneurophion Ashmead, 1900

         Cymatoneura Kriechbaumer, 1901

         Pterospilus Kriechbaumer, 1901; preocc. by Rondani, 1856

         Trispilus Kriechbaumer, 1901

         Metophion Szépligeti, 1905

         Ceratospilus Szépligeti, 1905

         Atoponeura Szépligeti, 1905

         Ophiomorpha Szépligeti, 1905; preocc. by Nilsson, 1836

         Cryptocampus Brèthes, 1909

         Eremotyloides Perkins, 1915

         Amesospilus Enderlein, 1918

         Schizospilus Seyrig, 1936

      5. Pycnophion Ashmead, 1900; Hawaii
2. Eremotylus genus-group
   1. Eremotylus Förster, 1869; Holarctic

      Camptoneura Kriechbaumer, 1901; preocc. by Agassiz, 1846, and by Fieber, 1858

      Genophion Felt, 1903 (Gauld, 1979)

      Camptoneuroides Strand, 1928

      Clistorapha Cushman, 1947 (Gauld, 1979)

      Boethoneura Cushman, 1947 (Gauld, 1979)

      Chilophion Cushman, 1947 (Gauld, 1979)

      Chlorophion Townes, 1971 (Gauld, 1979)

   2. Hellwigiella Szépligeti, 1905; Palearctic
   3. Trophophion Cushman, 1947; Nearctic
3. Ophion genus-group
   1. Afrophion Gauld, 1979; Ethiopian
   2. Agathophiona Westwood, 1882; Neotropical
   3. Alophophion Cushman, 1947; Neotropical
   4. Ophion Fabricius, 1798; worldwide except Ethiopian and Australian

      Paniscus Schrank, 1802

      Psylonychia Szépligeti, 1905, nomen nudum

      Stenophthalmus Szépligeti, 1905; preocc. by Becker, 1903

      Pachyprotoma Kohl, 1906

      Australophion Morley, 1912

      Neophion Morley, 1912

      Apatophion Shestakov, 1926 (Gauld, 1979)

      Platophion Hellén, 1926

      Potophion Cushman, 1947

      Psylonychia Cushman, 1947

      Apomesus Townes, 1971 (Gauld, 1979)

      Mecetron Townes, 1971 (Gauld, 1979)

   5. Rhopalophion Seyrig, 1935; Ethiopian
   6. Sclerophion Gauld, 1979; Oriental, Palearctic
   7. Xylophion Gauld, 1979; Australian
4. Sicophion genus-group
   1. Janzophion Gauld, 1985; Neotropical
   2. Riekophion Gauld, 1977a; Australian
   3. Sicophion Gauld, 1979; Neotropical
   4. Thyreodon genus-group
   5. Barytatocephalus Schulz, 1895; Palearctic

      Barycephalus Brauns, 1895; preocc. by Guenther, 1860

   6. Dictyonotus Kriechbaumer, 1894; Ethiopian, Oriental, Palearctic

      Aglaophion Cameron, 1903

      Ophionopsis Tosquinet, 1903 (Gauld, 1985)

      Hybopleurax Enderlein, 1912

      Hypselogasterina Enderlein, 1914 (Gauld, 1985)

      Coracophion Shestakov, 1926

   7. Euryophion Cameron, 1906; Ethiopian, Oriental

      Eurycamptus Morley, 1912

      Thoracophion Roman, 1943

      Primophion Townes, 1971 (Gauld & Mitchell, 1981)

      Rictophion Townes, 1971 (Gauld, 1985)

   8. Rhynchophion Enderlein, 1912; Nearctic, Neotropical
   9. Thyreodon Brullé, 1846; Nearctic, Neotropical

      Athyreodon Ashmead, 1900

      Tipulophion Kriechbaumer, 1901

      Macrophion Szépligeti, 1905

      Oleter Shestakov, 1926